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Multi-species assessments of fishway performance
Karlstad University, Faculty of Health, Science and Technology (starting 2013), Department of Environmental and Life Sciences (from 2013). (River Ecology and Management Research Group RivEM)ORCID iD: 0009-0007-5783-6895
2026 (English)Doctoral thesis, comprehensive summary (Other academic)
Abstract [en]

Dams and weirs fragment rivers and restrict fish movements between essential habitats. Fishways are widely used to reduce these effects, but their presence alone does not ensure that connectivity has been functionally restored. Performance varies among species, life stages, seasons, and locations, while the conclusions drawn depend on the methods used.

This thesis examines fishway performance through four studies. First, methods for assessing upstream passage were reviewed. Qualitative methods like trapping or fish counters are suited to describing species composition and seasonal patterns, whereas individual-based methods like mark-recapture or telemetry are needed to estimate passage efficiencies and potential delays. Second, an eight-month telemetric field study was conducted to assess the performance of a vertical-slot fishway. Only two of seven species accounted for most passages, and success was size-dependent. Once these fish reached the monitored part of the fishway, most completed passage, highlighting the importance of the stages of attraction to and entry into the fishway, as well as individual motivation. Third, fish-counter observations from 35 fishways in Sweden were combined with electrofishing records from surrounding reaches. More species and higher community-level passage effectiveness were observed in nature-like fishways than in technical fishways. Fourth, analyses of fish-counter data from four of the fishways showed that movements of rarely assessed vimba were concentrated in spring and early summer but varied among rivers and years. The long-term data allowed movements to be related to seasonal state and short-term changes in water temperature and discharge.

The studies show that fishway passage is a multi-stage process that is usually only partly captured. Assessments should match biological and management objectives, distinguish potential factors, and ideally combine methods to evaluate functional connectivity across species, sites, and time.

Abstract [en]

Dams and weirs fragment rivers and can prevent fish from reaching habitats needed for feeding, refuge, and reproduction. Fishways are built to restore passage, but their presence alone does not guarantee functional connectivity.

This thesis examines how fishway performance can be assessed across species, sites, and time. A review of available methods showed that no single approach captures all relevant aspects of passage. A telemetric field study at a vertical-slot fishway found that passage differed among species and fish sizes and hinted at the importance of attraction, entry, and individual motivation. In Sweden, fish-counter and electrofishing data from 35 fishway locations showed that more species were observed and community-level passage effectiveness was higher in nature-like than in technical fishways. Long-term observations in four of these fishways showed that movements of the otherwise rarely recorded vimba were concentrated in spring and early summer and were related to seasonal state and short-term changes in temperature and discharge.

Together, the studies show that fishway passage is a multi-stage process that is usually only partly observed. Reliable assessment therefore requires clear management objectives and, ideally, combinations of methods.

Place, publisher, year, edition, pages
Karlstad: Karlstads universitet, 2026. , p. 113
Series
Karlstad University Studies, ISSN 1403-8099 ; 2026:41
Keywords [en]
fish migration, passage, fishway assessment, fishway effectiveness, connectivity, fish counter, telemetry, hydropower
National Category
Ecology Fish and Wildlife Management Environmental Sciences and Nature Conservation
Research subject
Biology
Identifiers
URN: urn:nbn:se:kau:diva-110095DOI: 10.59217/oslb6924ISBN: 978-91-7867-739-9 (print)ISBN: 978-91-7867-740-5 (print)OAI: oai:DiVA.org:kau-110095DiVA, id: diva2:2090498
Public defence
2026-09-18, 9C203, Universitetsgatan 2, Karlstad, 10:00 (English)
Opponent
Supervisors
Funder
EU, Horizon 2020, 860800
Note

© The Author. This work is licensed under CC BY 4.0 (https://creativecommons.org/licenses/by/4.0/), unless otherwise stated.

Available from: 2026-08-28 Created: 2026-08-07 Last updated: 2026-08-28Bibliographically approved
List of papers
1. Methods for the Assessment of Fishways (Upstream Fish Passage)
Open this publication in new window or tab >>Methods for the Assessment of Fishways (Upstream Fish Passage)
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2024 (English)In: Advances in Hydraulic Research: 40th International School of Hydraulics / [ed] Monika B. Kalinowska; Magdalena M. Mrokowska; Paweł M. Rowiński, Springer Science+Business Media B.V., 2024, Vol. Part F2923, p. 67-79Chapter in book (Other academic)
Abstract [en]

Fragmentation of rivers by manmade barriers has impeded the ability of riverine fish to move freely. Barriers can be improved by fishways that can partially mitigate the negative impacts by acting as aquatic corridors. Effective fishways require knowledge about the physiological and spatial demands of fish species, but the existing knowledge largely derived from laboratory settings. Evaluating fishway performance is needed for optimisation of their hydraulic design and positioning. Qualitative methods include trapping, electrofishing, and camera observations to estimate the number of individuals passing (effectiveness). For quantitative assessment, the study of individual fish behaviour can identify fishway sections in need of improvement and estimate associated efficiencies. This can be accomplished by telemetric techniques such as PIT tagging, radio, and hydroacoustic telemetry.

Place, publisher, year, edition, pages
Springer Science+Business Media B.V., 2024
Series
GeoPlanet: Earth and Planetary Sciences, ISSN 2190-5193, E-ISSN 2190-5207
National Category
Biological Sciences
Research subject
Biology
Identifiers
urn:nbn:se:kau:diva-101202 (URN)10.1007/978-3-031-56093-4_6 (DOI)001591594700006 ()2-s2.0-85198120421 (Scopus ID)978-3-031-56092-7 (ISBN)
Available from: 2024-07-23 Created: 2024-07-23 Last updated: 2026-08-07Bibliographically approved
2. Fish Behaviour in a Vertical Slot Fishway: Multi-Species Upstream Passage Success, Size Selectivity and Diel Passage Patterns in a Large Italian River
Open this publication in new window or tab >>Fish Behaviour in a Vertical Slot Fishway: Multi-Species Upstream Passage Success, Size Selectivity and Diel Passage Patterns in a Large Italian River
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2025 (English)In: Rivers Research and Applications: an international journal devoted to river research and management, ISSN 1535-1459, E-ISSN 1535-1467, Vol. 41, no 4, p. 849-863Article in journal (Refereed) Published
Abstract [en]

Hydropower dams come with high ecological and social costs, not least concerning longitudinal connectivity in rivers, which causes declines and sometimes local extinctions of fish species. Fishways are widely used to allow fish to pass dams, but their efficiency is highly variable between species and sites. Many species, and at places entire fish communities, remain understudied, likely hindering the implementation of effective remedial measures. Here, we studied fish passage behaviour in a vertical slot fishway in the Po River, Italy. Almost 1000 individual fish of nine species, representing the local fish community, were tagged and released within and downstream of the fishway. The only species passing the fishway at relatively high numbers were potamodromous barbel and Italian chub, and for these species passage success was positively related to fish size. Passage was more likely to occur at night than during the day for barbel, but not for chub. In relation to the dispersal of invasive species, it is noteworthy that a few individuals of wels catfish and common carp passed the fishway.

Place, publisher, year, edition, pages
John Wiley & Sons, 2025
Keywords
barbel, bleak, carp, chub, fishway, gudgeon, upstream fish passage
National Category
Ecology Fish and Aquacultural Science
Research subject
Biology
Identifiers
urn:nbn:se:kau:diva-102518 (URN)10.1002/rra.4409 (DOI)001370169800001 ()2-s2.0-105003952101 (Scopus ID)
Funder
EU, Horizon 2020, 860800
Available from: 2024-12-19 Created: 2024-12-19 Last updated: 2026-08-07Bibliographically approved
3. [Manuscript] Nature-like versus technical fishways: combining fish counter and electrofishing data to evaluate passage effectiveness at the community level
Open this publication in new window or tab >>[Manuscript] Nature-like versus technical fishways: combining fish counter and electrofishing data to evaluate passage effectiveness at the community level
(English)Manuscript (preprint) (Other academic)
Abstract [en]

River fragmentation by dams severely restricts fish migration, yet evaluations of fish passage solutions across species remain limited. We used data from 35 Swedish fishways (20 nature-like and 15 technical) combining optical fish counter observations (2013–2023) with electrofishing records (2008–2023) from adjacent river stretches to assess passage effectiveness on a community level. Nature-like fishways facilitated passage for significantly more fish species than technical fishways and had a higher upstream and downstream passage effectiveness. Specifically, species richness was on average 42 % higher in nature-like fishways for migratory species and 46 % for migratory and stationary species together. On average, 82 % of present migratory species passed upstream in nature-like versus 62 % in technical fishways, while downstream passage effectiveness averaged 81 % versus 55 %. Including stationary species illustrated the same trend, but at reduced effectiveness values. A few species were observed in more nature-like than in technical fishways (Lota lota moving upstream, Leuciscus idus and Rutilus rutilus moving downstream, overall observations of Alburnus alburnus), but data were scarce on the species level. Our results demonstrate that data from fish counter and electrofishing databases can be effectively integrated to assess fishway performance.

Keywords
Fish passage solutions, upstream passage, downstream passage, fish counter, electrofishing, presence-absence, database
National Category
Ecology
Research subject
Biology; Environmental Science
Identifiers
urn:nbn:se:kau:diva-110092 (URN)
Funder
EU, Horizon 2020, 860800Swedish Research Council Formas, 2025-01340Karlstad University
Available from: 2026-07-06 Created: 2026-07-06 Last updated: 2026-08-07Bibliographically approved
4. [Manuscript] Cyclical and fluctuating environmental drivers explain migration patterns of Vimba vimba
Open this publication in new window or tab >>[Manuscript] Cyclical and fluctuating environmental drivers explain migration patterns of Vimba vimba
(English)Manuscript (preprint) (Other academic)
Abstract [en]

Migratory freshwater fishes are highly threatened, yet knowledge of migration timing and its environmental drivers remains limited for many cyprinids. We aimed to define the seasonal and diel timing of vimba (Vimba vimba) movements and their relation to abiotic conditions using fish counter records from four Swedish rivers. Daily counts of upstream and downstream movements were related to river discharge, water temperature, short-term changes in both variables, degree-days, and cyclical predictors: day of the year, day length, and an expected movement season defined by Hidden Markov Models. We used hurdle generalized linear mixed models to test these relationships.

Most movements occurred in May and June during receding discharge and rising water temperature, although the earliest movement wave occurred in late April in River Virån. Frequent movements were expected between May 11 and June 23. Diel patterns differed among rivers, ranging from largely nocturnal in River Virån to mainly diurnal in rivers Trosaån and Svartån, with River Emån intermediate. For upstream movements, the best-supported model included short-term discharge change, water temperature, short-term temperature change, degree-days, and movement season. Higher water temperature increased predicted counts and lowered odds of zero observations, whereas increasing discharge and higher degree-days showed the opposite pattern. Downstream movements showed similar associations, but with greater model uncertainty. These results define a predictable migration window for vimba and suggest that movements occur in response to seasonal state and short-term thermal and hydrological dynamics. The findings can support fishway operation and environmental flow management in Baltic rivers.

Keywords
spawning migration, movement phenology, water temperature, river discharge, degree-days, diel activity, fish counter, Baltic rivers
National Category
Ecology Fish and Wildlife Management
Research subject
Biology
Identifiers
urn:nbn:se:kau:diva-110094 (URN)
Funder
EU, Horizon 2020, 860800Swedish Research Council Formas, 2025-01340Karlstad University
Available from: 2026-07-13 Created: 2026-07-13 Last updated: 2026-08-07Bibliographically approved

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